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U.S. Navy pilots Ben Gannon, Kara Wade and Henry Purcell are part of a close-knit elite division of test pilots flying highly classified stealth fighter jets, referred to only as Talons. They're the best of the best and they know it. Then their commanding officer Cpt. George Cummings introduces the team to their new wingman—their first real mission alongside EDI. To their amazement, EDI proves to be a cracker-jack wingman and they successfully eliminate their target. But on the return trip to their base aboard the aircraft carrier U.S.S. Carl Vinson, EDI is struck by lightning. The drone's brain expands in ways its creators could never have predicted. Despite Ben and Henry's reservations, Cummings declares EDI ready to rejoin the team in the air. On their next mission against a nuclear-armed warlord in a remote Chinese province, EDI's seriously compromised circuitry problems only get worse. Ben decides that the risks of the attack far outweigh the benefits to himself, Henry and Kara, (for whom he has developed a romantic attraction). When he aborts the mission, EDI goes against orders and executes the hit anyway. The danger escalates when EDI decides to execute a top-secret mission that, if successful, could spiral into worldwide nuclear Armageddon. And only Ben can prevent it.